glucose
From The Oxford Companion to Beer
is the building block of starch and as such is the primary source of fermentable material in wort and, subsequently, of alcohol in beer. Chemically, glucose is a polyhydroxy aldehyde, with a chemical formula of C6H12O6.
In starch, glucose occurs either as long straight chains (amylose) or as branched chains (amylopectin), in which the glucose molecules are joined by either alpha 1,4 or alpha 1,6 linkages. Brewing yeasts are incapable of metabolizing starches. These must first be enzymatically degraded during malting and mashing into mostly maltose, which is made up of two glucose molecules joined with alpha 1,4 linkages, and maltotriose, which consists of three-linked glucose units. Smaller amounts of glucose are also produced during mashing, particularly toward the end of the process. Typically, all-malt wort contains about 7% glucose, 45% maltose, and 20% maltotriose. The remaining roughly 25% of carbohydrates are sucrose and fructose, as well as unfermentable oligosaccharides and dextrins. During wort boiling some of the glucose reacts with amino compounds (mainly from amino acids) to create colored compounds, which both increase beer color and can impart toffee or caramel-like flavors.
Bibliography
This definition is from The Oxford Companion to Beer, edited by Garrett Oliver. © Oxford University Press 2012.